Off-season flowering in red pitaya (Hylocereus sp.), a long-day plant, can be achieved using night-breaking (NB) treatment. The ovules are circinotropous, bitegmic, crassinucelate, and have long funiculus as in many other Cactaceae. The stigma has 9-10 lobes with a secretory epidermis. The nectariferous region occurs on the inner surface of the hypanthium. The ovary has inverted vascular bundles in a similar pattern as in Pereskia. The gynoecium is syncarpous with 9-10 carpels, pluriovulate, and with parietal placentation. The wall of the young anther is formed by epidermis, endothecium, a middle layer, and binucleate secretory tapetum that eventuallyīecomes uninucleate. The androecium has many stamens with tetrasporangiate and bithecal anthers. Secretory cavities containing mucilage and calcium oxalate crystals occur throughout the floral All fl ower whorls have uniseriate epidermis. The fl ower is sessile and epigynous with a well-developed hypanthium. Flowers and fl ower buds were collected in Maringá, Paraná State, Brazil, fi xed, processed, and analyzed under light microscope and scanning electron microscope. The aim of this contribution was to analyze the morpho-anatomical fl oral structure of Epiphyllum phyllanthus (L.) Haw., a widely distributed species across South America, occurring in humid forests as an epiphyte.
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